CURRICULUM VITAE: Susan Patricia Harrison

Department of Environmental Science and Policy
(530) 752-7110,

Academic Postitions Grants
Awards (Post-PhD) Professional Service
Education University Service
Scientific Articles Public Service
Books & Book Chapters Invited Lectures
Book Reviews  


Research Associate, Center for Conservation Biology, Stanford (1986 - 1989)

Postdoctoral Fellow, Morrison Institute for Population and Resource Studies, Stanford (1989)

Postdoctoral Researcher, Centre for Population Biology, Imperial College at Silwood Park, England (1990 - 1991)

Assistant Professor, Division of Environmental Studies, U.C. Davis (1991 - 1996)

Associate Professor, Division of Environmental Studies, U.C. Davis (1996 - 1998)

Professor, Department of Environmental Science and Policy, U.C. Davis (1998 -)

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International Recognition of Professional Excellence Award (International Ecology Institute, Oldendorf, Germany), 1996
Honorary Fellow, California Academy of Sciences, 2004-
Fellow, Ecological Society of America, 2013-


B.S. (Zoology) 1983, University of California, Davis
M.S. (Ecology) 1986, University of California, Davis
Ph.D. (Biology) 1989, Stanford University

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Spasojevic, M., J. B. Grace, E. I. Damschen and S. Harrison 2014. Functional diversity supports the physiological tolerance hypothesis for plant species richness along climatic gradients. Journal of Ecology (in press).

Eskelinen, A. M. and S. Harrison 2013. Exotic plant invasions under enhanced rainfall are constrained by soil nutrients and competition. Ecology (in press).

Fernandez, B. M. and S. Harrison 2013. Effects of experimental water addition depend on grassland community characteristics. Plant Ecology (in press).

Fernandez, B. M., B. L. Anacker, S. Harrison, and H.D. Safford 2013. Climate interacts with soil to produce beta diversity in the Californian flora. Ecology (in press).

Hulshof, C., B. Enquist, M. Spasojevic, C. Violle, E. Damschen and S. Harrison 2013. Intraspecific and interspecific variation in specific leaf area reveal the importance of abiotic and biotic drivers of species diversity across elevation and latitude. Journal of Vegetation Science (in press).

Cornell, H. V. and S. P. Harrison 2013. Regional effects as important determinants of local diversity in both marine and terrestrial systems. Oikos (in press).

Bieger, A., N. Rajakaruna and S. Harrison 2013. Little evidence for local adaptation to soils or microclimate in the post-fire recruitment of three Californian shrubs. Plant Ecology and Diversity (in press).

Eskelinen, A., S. Harrison and M.. Tuomi 2013. Plant traits mediate consumer and nutrient control on plant community productivity and diversity. Ecology 93: 2705-2718

Spasojevic, M., E. Damschen and S. Harrison 2013. Patterns of seed dispersal syndromes on serpentine soils: examining the roles of habitat patchiness, soil infertility, and correlated functional traits. Plant Ecology & Diversity DOI:10.1080/17550874.2012.678506

Damschen, E. I., S. Harrison, D. D. Ackerly, B. M. Fernandez-Going and B. L. Anacker 2012. Endemic plant communities on special soils: Early victims or hardy survivors of climate change? Journal of Ecology (in press). DOI: 10.1111/j.1365-2745.2012.01986.x

Fernandez-Going, B. M., B. L. Anacker, and S. Harrison 2012. Temporal variability in California grasslands: soil type and species functional traits mediate response to precipitation. Ecology 93: 2104-2114.

Anacker, B. L. and S. Harrison 2012. Historical and ecological controls on phylogenetic diversity in Californian plant communities. American Naturalist 180: 257-269

Anacker, B. L., N. Rajakaruna, D. D. Ackerly, M. C. Vasey, S. Harrison, and J. E. Keeley. 2012. Ecological strategies in California chaparral: Interacting effects of soils, climate, and fire on specific leaf area. Plant Ecology & Diversity 4: 179-188.

Hawkins, B. A., C. M. McCain, T. J. Davies, L. B. Buckley, B. Anacker, H. V. Cornell, E. I. Damschen, J.-A. Grytnes, S. Harrison, R. D. Holt, N. J. B. Kraft and P. R. Stephens 2012. Different evolutionary histories underlie congruent species richness gradients of birds and mammals. Journal of Biogeography 39: 825-841.

Stegen, J. C., A. L. Freestone, T. O. Crist, M. J. Anderson, J. M. Chase, L. S. Comita, H.V. Cornell, K. F. Davies, S. P. Harrison, A. H. Hurlbert, B.D. Inouye, N. J. B. Kraft, J. A. Myers, N. J. Sanders, N. G. Swenson, M. Vellend. 2011. Stochastic and deterministic drivers of spatial and temporal turnover in breeding bird communities. Global Ecology and Biogeography (in press).

Anacker, B. L. and S. Harrison 2012. Climate and the evolution of serpentine endemism in California. Evolutionary Ecology 26: 1011-1023.

Kraft, N. J. B., L. S. Comita, J. M. Chase, N. J. Sanders, N. G. Swenson, T. O. Crist, J. C. Stegen, M. Vellend, B. Boyle, M. J. Anderson, H.V. Cornell, K. F. Davies, A. L. Freestone, B.D. Inouye, S. P. Harrison, and J. A. Myers 2011. Disentangling the drivers of beta-diversity along latitudinal and elevational gradients. Science 333: 1755-1758.

Moore, K. A., S. Harrison and S. C. Elmendorf 2011. Can spatial isolation help predict dispersal-limited sites for native species restoration? Ecological Applications. Ecological Applications 21: 2119-2128.

Harrison, S., M. Vellend and E. I. Damschen 2011. ‘Structured’ beta diversity increases with climatic productivity in a classic dataset. Ecosphere 2:article11.

Harrison, S., E. I. Damschen and J. B. Grace 2010. Ecological contingency in the effects of climate change on forest herbs. Proceedings of the National Academy of Sciences (USA), 107: 19362-19367.

Anacker, B. L., J. Whittall, E. Goldberg, and S. Harrison 2011. Origins and consequences of serpentine endemism in the California flora. Evolution (in press).

Anderson, M. J., T. O. Crist, J. M. Chase, M. Vellend, B. D. Inouye, A. L. Freestone, N. J. Sanders, H. V. Cornell, L. S. Comita, K. F. Davies, S. P. Harrison, N. J. B. Kraft, J.C. Stegen, and N. G. Swenson 2011. Navigating the multiple meanings of beta diversity: a roadmap for the practicing ecologist. Ecology Letters (in press).

Elmendorf, S. and S. Harrison. Is plant community diversity regulated over time? Contrasting results from experiments and long-term observations. Ecology 92: 602-609.

Grace, J. B., S.Harrison and E. I., Damschen 2011. Local richness along gradients in the Siskiyou herb flora: R. H. Whittaker revisited. Ecology 92: 108-120.

Damschen E. I., S.Harrison and J. B. Grace, 2010. Climate change effects on an endemic-rich edaphic flora: resurveying Robert H. Whittaker's Siskiyou sites (Oregon, USA). Ecology 91: 3609-3619

Wiens, J., D. Ackerly, A. Allen, B. Anacker, L. Buckley, H. Cornell, E. Damschen, J. Davies, J.-A. Grytnes, S.Harrison, B. Hawkins, R. Holt, C. McCain, P. Stephens, 2011. Niche conservatism as an emerging principle in ecology and conservation biology. Ecology Letters 13: 1310-1324.

Buckley, L. B., T. J.Davies, D. D. Ackerly, N. J. B. Kraft, S. P. Harrison, B. L. Anacker, H. V. Cornell, E. I. Damschen, J.-A. Grytnes, B. A. Hawkins, C. M. McCain, P. R. Stephens0, and J. J. Wiens 2010. Phylogeny, niche conservatism, and the latitudinal diversity gradient in mammals. Proceedings of the Royal Society B 277: 2131-2138.

Harrison, S., H. V. Cornell and K. A. Moore 2010. Spatial niches and coexistence: testing theory with tarweeds. Ecology 91: 2141-2150.

Harrison, S., E. I . Damschen and B. M. Going 2009. Climate gradients, climate change, and special edaphic floras. Northeastern Naturalist 16: 121-130.

Elmendorf, S. and S. Harrison 2009. Temporal variability and nestedness in Californian grassland species composition. Ecology 90: 1492-1497.

Harrison, S. and H. V. Cornell, 2008. Toward a better understanding of the regional causes of local community richness. Ecology Letters 11: 1-11.

Harrison, S., J. H. Viers, J. H. Thorne, and J. B. Grace, 2008. Favorable environments and the persistence of naturally rare species. Conservation Letters 1: 65-74.

Harrison, S. and A. R. Kruckeberg 2008. Garden on the rocks. Natural History (May): 40-44.

Harrison, S. and J. B. Grace 2007. Biogeographic affinity helps explain the productivity-richness relationship at regional and local scales. American Naturalist 170: S5-S15.

Harrison, S. and H. V. Cornell 2007. Introduction: merging evolutionary and ecological approaches to understanding geographic gradients in species richness. American Naturalist 170: S1-S4.

Davies, K. F., S. Harrison, H. D. Safford, and J. H. Viers 2007. Productivity alters the scale dependence of the diversity-invasibility relationship. Ecology 88: 1940-1947.

Hugueny, B., H. V. Cornell and S. Harrison 2007. Simple metacommunity models predict the local-regional species richness relationship in a natural system. Ecology 88: 1696-1706.

Mittelbach, G., D. Schemske, H. Cornell, A. Allen, J. Brown, M. Bush, S. Harrison, A. Hurlbert, N. Knowlton, H. Lessios, C. McCain, A. McCune, L. McDade, M. McPeek, T. Near, T. Price, R. Ricklefs, D. Sax, D. Schluter, and J. Sobel 2007. Evolution and the latitudinal diversity gradient: speciation, extinction, and biogeography. Ecology Letters 10:315-331.

Grace, J. B., H. D. Safford, and S. Harrison 2007. Large-scale causes of variation in the serpentine vegetation of California. Plant and Soil 293: 121-132.

Harrison, S., H. D. Safford, J. B. Grace, J. H. Viers and K. F. Davies, 2006. Regional and local species richness in an insular environment: serpentine plants in California. Ecological Monographs 76: 41-56.

Harrison, S., K. F. Davies, J. B. Grace, H. D. Safford, and J. H. Viers 2006. Exotic invasion in a diversity hotspot: disentangling the direct and indirect relationships of exotic cover to native richness in the Californian serpentine flora. Ecology 87: 695-703.

Harrison, S., K. F. Davies, H. D. Safford, and J. H. Viers 2006. Beta diversity and the scale-dependence of the productivity-diversity relationship: a test in the Californian serpentine flora. Journal of Ecology 94: 110-117

Freestone, A. L. and S. Harrison 2006. Regional enrichment of local assemblages is robust to variation in local productivity, abiotic gradients, and heterogeneity. Ecology Letters 9: 95-102.

Melbourne, B., H. Cornell, K. Davies, C. Dugaw, S. Elmendorf, A. Freestone, R. Hall, S. Harrison, A. Hastings, M. Holland, M. Holyoak, J. Lambrinos, K. Moore, and H. Yokomizo 2006. Invasion in a heterogeneous world: resistance, coexistence or hostile takeover? Ecology Letters 10: 77-94.

Batten, K. M., K. M. Scow, K. F. Davies, and S. Harrison, 2006. Two invasive plants alter soil microbial community composition in serpentine grasslands. Biological Invasions 8: 217-230.

Davies, K. F., P. Chesson, S. Harrison, B. D. Inouye, B. A. Melbourne, and K. J. Rice, 2005. Spatial heterogeneity explains the scale dependence of the native-exotic diversity relationship. Ecology 86: 1602-1610.

Gelbard, J. L. and S. Harrison, 2005. Invasibility of roadless grasslands: an experimental study of yellow starthistle. Ecological Applications 15: 1570-1580.

Harrison, S., A. Hastings and D. R. Strong, 2005. Spatial and temporal dynamics of insect outbreaks in a complex multitrophic system: tussock moths, ghost moths, and their natural enemies on bush lupines. Annales Zoologici Fennici 42: 409-419

McKay, J. K., C. E. Christian, S. Harrison and K. J. Rice 2005. “How local is local?” A review of practical and conceptual issues in the genetics of restoration. Restoration Ecology 13: 432-440.

Safford, H. D., J. H. Viers and S. Harrison, 2005. Serpentine endemism in the Calfornia flora: a database of serpentine affinity. Madroño 52: 222-257.

Harrison, S. and H. D. Safford, 2004. Regional and local diversity in the serpentine endemic flora of California: preliminary results of a geographic study. Pages 183-190 in R.

S. Boyd, A. J. M. Baker, and J. Proctor (eds.), Ultramafic rocks: their soils, vegetation, and fauna. (Proceedings of the Fourth International Conference on Serpentine Ecology). Science Reviews, St. Albans, UK.

Safford, H. D. and S. Harrison, 2004. The effects of fire and grazing on serpentine versus nonserpentine grassland and chaparral. Pages 315-322 in R. S. Boyd, A. J. M. Baker, and J. Proctor (eds.), Ultramafic rocks: their soils, vegetation, and fauna. (Proceedings of the Fourth International Conference on Serpentine Ecology). Science Reviews, St. Albans, UK.

Harrison, S., H. D. Safford, and J. Wakabayashi, 2004. Does the age of exposure of serpentine explain variation in endemic plant diversity in California? International Geology Review 46: 235-242. (Reprinted in: Serpentine and serpentinites: mineralogy, petrology, geochemistry, ecology, geophysics, and tectonics [the Coleman Volume], International Geology Series of the GSA, Bellwether Publishers, UK).

Harrison, S. 2004. Biodiversity and wilderness. Journal of Land, Resources, & Environmental Law, fall 2004. (Proceedings from the Ninth Annual Wallace Stegner Center Symposium, S. J. Quinney School of Law, Univ. of Utah, April 2004.)

Hastings, A., K. Cuddington, K. Davies, C. Dugaw, S. Elmendorf, A. Freestone, S. Harrison, M. Holland, J. Lambrinos, U. Malvadkar, B. Melbourne, K. Moore, C. Taylor and D. Thomson, 2004. The spatial spread of invasions: new developments in theory and evidence. Ecology Letters 8: 91-101.

Safford, H. D. and S. Harrison, 2004. Fire effects on plant diversity in serpentine versus sandstone chaparral. Ecology 85: 539-548.

Harrison, S., B. D. Inouye and H. D. Safford, 2003. Ecological heterogeneity in the effects of grazing and fire on grassland diversity. Conservation Biology 17: 837-845.

Gelbard, J. L. and S. Harrison, 2003. Roadless habitats as refuges for native plant diversity in California grassland landscapes. Ecological Applications 13: 404-415.

Umbanhowar, J., J. Maron and S. Harrison, 2003. Density-dependent foraging behaviors in a parasitoid lead to density-dependent parasitism of its host. Oecologia 137:123-130.

Williamson, J. N. and S. Harrison 2002. Biotic and abiotic limits to the spread of exotic revegetation species in oak woodland and serpentine habitats. Ecological Applications 12: 40-51.

Jurjavcic, N. L., S. Harrison and A. T. Wolf, 2002. Abiotic stress, competition, and the distribution of the native annual grass Vulpia microstachys in a mosaic environment. Oecologia 130: 555-562.

Harrison, S. and B. D. Inouye, 2002. High beta diversity in the flora of Californian serpentine "islands". Biodiversity and Conservation 11: 1869-1876.

Bruna, E. M., O. Nardy, S. Y. Strauss, and S. Harrison, 2002. Experimental assessment of Heliconia acuminata growth in a fragmented Amazonian landscape. Journal of Ecology 90: 639-649.

Harrison, S., C. Hohn and S. Ratay, 2002. Distribution of exotic plants along roads in a peninsular nature reserve. Biological Invasions 4: 425-430.

Safford, H. and S. Harrison, 2001. Ungrazed roadside verges in a grazed landscape: interactive effects of grazing, invasion and substrate on grassland diversity. Ecological Applications 11: 1112-1122.

Maron, J., S. Harrison and M. Greaves, 2001. Origin of an insect outbreak: escape in space or time from natural enemies? Oecologia 126: 595-602.

McCann, K. S., A. M. Hastings, S. Harrison and W. G. Wilson, 2000. Population outbreaks in a discrete world. Theoretical Population Biology 57: 97-108.

Wolf, A. T., S. Harrison and J. L. Hamrick, 2000. The influence of habitat patchiness on genetic diversity and spatial structure of a serpentine endemic plant. Conservation Biology 14: 454-463.

Harrison, S., J. Maron and G. Huxel, 2000. Regional turnover and fluctuation in populations of five plants confined to serpentine seeps. Conservation Biology 14: 769-779.

Harrison, S., J. L. Viers and J. F. Quinn, 2000. Climatic and spatial patterns of diversity in the serpentine plants of California. Diversity and Distributions 6: 153-161.

Wolf, A. T. and S. Harrison 2001. Natural habitat patchiness affects reproductive success of serpentine morning glory (Calystegia collina, Convolvulaceae) in northern California. Conservation Biology 15: 111-121.

Harrison, S., K. Rice and J. Maron, 2001. Habitat patchiness promotes invasions by alien grasses on serpentine soil. Biological Conservation 100: 45-53.

Harrison, S. 1999. Local and regional diversity in a patchy landscape: native, alien and endemic herbs on serpentine soils. Ecology 80: 70-80.

Wilson, W. G., S. Harrison, A. Hastings and K. McCann 1999. Exploring spatial pattern formation in models of tussock moth populations. Journal of Animal Ecology 68: 94-107.

Wolf, A. T., P. A. Brodmann and S. Harrison 1999. Distribution of the rare serpentine sunflower (Helianthus exilis Gray, Asteraceae): the roles of habitat availability, dispersal limitation and species interactions. Oikos 84: 69-76.

Marsh, D. M., E. H. Fegraus, and S. Harrison 1999. Effects of breeding pond isolation on the spatial and temporal dynamics of pond use in the tungara frog, Physaelaemus pustulosus. Journal of Animal Ecology 68: 584-594.

Harrison, S. and E. M. Bruna 1999. Habitat fragmentation and large-scale conservation: what do we know for sure? Ecography 22: 1-8.

Harrison, S. 1999. Native and alien species diversity at the local and regional scales in a grazed Californian grassland. Oecologia 121: 99-106.

Harrison, S. 1997. How natural habitat patchiness affects the distribution of diversity in Californian serpentine chaparral. Ecology 78: 1898-1906.

Harrison, S. 1997. Persistent, localized outbreaks in the western tussock moth (Orgyia vetusta): the roles of resources, predation and poor dispersal. Ecological Entomology 22: 158-166.

Hastings, A., Harrison, S. and McCann, K. S. 1997. Unexpected spatial patterns in an insect outbreak match the predictions of a predator diffusion model. Proceedings of the Royal Society of London, Series B 264: 1837-1840.

Maron, J. L. and S. Harrison, 1997. Spatial pattern formation in an insect host-parasitoid system. Science 278: 1619-1621.

Harrison, S. and A. M. Hastings, 1996. Genetic and evolutionary consequences of metapopulation structure. Trends in Ecology and Evolution 11: 180-183.

Harrison, S. and A. M. Hastings, 1996. How effective is interdemic selection? A reply to Goodnight. Trends in Ecology and Evolution 11: 299.

Brodmann, P. A., C. V. Wilcox and S. Harrison, 1996. Mobile parasitoids may restrict the spatial spread of an insect outbreak. Journal of Animal Ecology 66: 65-72.

Harrison, S., C.D. Thomas and T. M. Lewinsohn, 1995. Testing a metapopulation model of coexistence in the insect community on ragwort (Senecio jacobaea). American Naturalist 145: 546-562.

Harrison, S. and C. Wilcox, 1995. Evidence that predator satiation may restrict the spatial spread of a tussock moth (Orgyia vetusta) outbreak. Oecologia 101: 309-316.

Harrison, S. 1995. Lack of strong induced or maternal effects in tussock moths (Orgyia vetusta) on bush lupine (Lupinus arboreus). Oecologia 103: 343-348.

Harrison, S. and J. L. Maron, 1995. Impacts of defoliation by tussock moths (Orgyia vetusta) on bush lupine (Lupinus arboreus). Ecological Entomology 20: 223-229.

Strong, D. R., J. L. Maron, P. G. Connors, A. Whipple, S. Harrison and R. L. Jefferies, 1995. High mortality, fluctuation in numbers, and heavy subterranean insect herbivory in bush lupine, Lupinus arboreus. Oecologia 104: 85-92.

Hastings, A. and S. Harrison, 1994. Metapopulation dynamics and genetics. Annual Review of Ecology and Systematics 25: 167-188.

Harrison, S. 1994. Resources and dispersal as factors limiting a population of the tussock moth (Orgyia vetusta), a flightless defoliator. Oecologia 99: 27-34.

Harrison, S., A. Stahl and D. F. Doak 1993. Spatial models and spotted owls: exploring some biological issues behind recent events. Conservation Biology 7: 950-953. (Reprinted in “Readings In Conservation Biology”, Vols I. To Preserve Biodiversity –An Overview; II. Wildlife and Forests; and III. The Landscape Perspective; ed. D. Ehrenfeld. Blackwell Scientific, Oxford, England.

Thomas, C.D. and S. Harrison 1992. Spatial dynamics of a patchily-distributed butterfly species. Journal of Animal Ecology 61: 437-446.

Harrison, S., J.F. Baughman, J.F. Quinn, P.R. Ehrlich and D.D. Murphy 1991. Estimating the effects of sampling on two butterfly populations. American Naturalist 137: 227-243.

Harrison, S. 1991. Population growth, land use and deforestation in Costa Rica, 1950-1984. Interciencia 16: 83-94.

Harrison, S. and C. D. Thomas 1991. Patchiness and spatial pattern in the insect community on ragwort (Senecio jacobaea). Oikos 62: 5-12.

Harrison, S., S. J. Ross and J. H. Lawton 1992. Beta diversity on geographic gradients in Britain. Journal of Animal Ecology 61: 141-148.

Harrison, S. 1989. Long-distance dispersal and colonization in the bay checkerspot butterfly, Euphydryas editha bayensis. Ecology 70: 1236-1243.

Harrison, S. and J.F. Quinn 1989. Correlated environments and the persistence of metapopulations. Oikos 56: 293-298.

Harrison, S., D.D. Murphy and P.R. Ehrlich 1988. Distribution of the bay checkerspot butterfly, Euphydryas editha bayensis: evidence for a metapopulation model. American Naturalist 132: 360-382.

Quinn, J.F. and S. Harrison 1988. Effects of habitat fragmentation and isolation on species richness: evidence from biogeographic patterns. Oecologia 75: 132-140.

Harrison, S. and A.M. Shapiro 1988. Butterflies of Northern California serpentines. Fremontia 15: 7-20.

Harrison, S. 1988. Local extinction, metapopulations and endangered species. Endangered Species Update 5(5): 10.

Harrison, S. 1987. Treefall gaps versus forest understory as environments for a defoliating moth on a tropical forest shrub. Oecologia 72: 65-68.

Harrison, S. and R. Karban 1986a. Effects of an early-season folivorous moth on the success of a later-season species, mediated by a change in the quality of the shared host, Lupinus arboreus Sims. Oecologia 69: 354-359.

Harrison, S. and R. Karban 1986b. Behavioural response of spider mites (Tetranychus urticae) to induced resistance of cotton plants. Ecological Entomology 11: 181-188.

Quinn, J.F., C. van Riper, R. Karban, G. Robinson and S. Harrison 1985. Conservation strategies: island biogeography and the design of nature reserves. Park Science 6: 6-9.

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Harrison, S. 2013. Plant and Animal Endemism in California. University of California Press.

Harrison, S. and N. Rajakaruna 2011. Serpentine: evolution and ecology of a model system. University of California Press.

Alexander, E. A., R. G. Coleman, T. Keeler-Wolf and S. Harrison 2006. Serpentine Geoecology of Western North America. Oxford University Press. 512 pp.

Harrison, S. and J. H. Viers, 2006. Serpentine Grasslands. In: Ecology and Management of California Grasslands. M.R. Stromberg, C. d’Antonio and J. Corbin, eds. Island Press.

Harrison, S. and C. Ray, 2000. Plant population viability and metapopulation-level processes. In S. Beissinger and D. McCullough (eds.), Population Viability Analysis. University of Chicago Press.

Harrison, S. 2000. Population persistence and community diversity in a naturally patchy landscape: plants on serpentine soils. In M. Kato (ed.), The Biology of Biodiversity. Springer Verlag, Tokyo.

Harrison, S. 2000. A field guide to studying spatial pattern formation in host-parasitoid systems. In M. E. Hochberg and A. R. Ives (eds.), Parasitoid Population Biology. Princeton University Press.

Hoopes, M. F. and S. Harrison. 1998. Metapopulation, source-sink and disturbance dynamics. In: Conservation Science and Action, ed. W. J. Sutherland, pp. 135-151. Blackwell, Oxford, England.

Harrison, S. and A. D. Taylor, 1997. Empirical evidence for metapopulation dynamics. In: Metapopulation Dynamics: Ecology, Genetics and Evolution, eds. I. Hanski and M. E. Gilpin, pp. 27-42. Academic Press, New York, USA.

Harrison, S. 1997. Do taxa persist as metapopulations in evolutionary time? In: Biodiversity Dynamics: Turnover of Populations, Taxa, and Communities, eds. M. L. McKinney and J. A. Drake. Columbia University Press, New York, USA.

Cappuccino, N. and S. Harrison, 1997. Density-perturbation experiments for understanding population regulation.. In: Frontiers in Population Ecology (Proceedings of the Nicholson Centenary Symposium), ed. A. Sheppard. CSIRO, Canberra, Australia.

Gutierrez, R. J. and S. Harrison, 1996. Applications of metapopulation theory to spotted owl management: a history and critique. In: Metapopulations and Wildlife Conservation Management, ed. D. McCullough, pp. 167-186. Island Press, Covelo, CA.

Harrison, S. and N. Cappuccino, 1995. Experimental approaches to understanding population regulation. In: Population Dynamics: New Approaches and Synthesis, eds. N. Cappuccino and P. W. Price. Academic Press, New York, USA.

Harrison, S. 1994. Metapopulations and conservation. In: Large-Scale Ecology and Conservation Biology. eds. P. J. Edwards, N. R. Webb and R. M. May, pp. 111-128. Blackwell Scientific, Oxford, England.

Harrison, S. and L. Fahrig, 1994. Landscape structure and population survival. In: Mosaic Landscapes and Ecological Processes, eds. L. Hansson, L. Fahrig and G. Merriam, pp. 293-308. Chapman and Hall, New York, USA.

Harrison, S. 1992. Species diversity, spatial scale and global change. In: Biotic Interactions and Global Change, eds. P. Kareiva, R. Huey and J. Kingsolver, pp. 388-401. Sinauer Press, Sunderland, MA, USA.

Harrison, S. 1991. Local extinction and metapopulation persistence: an empirical evaluation. Biological Journal of the Linnean Society 42: 73-88. (Reprinted in Metapopulation Dynamics: Empirical and Theoretical Investigations, eds. M. Gilpin and I Hanski, Academic Press, London.)

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Harrison, S. 2006. Book review. The shrinking Earth: ecological consequences of habitat loss, by I. Hanski. American Scientist 94: 82-83.

Harrison, S. 2003. Book review: Geology and plant life: the effects of landforms and rock types on plants, by A. R. Kruckeberg. Quarterly Review of Biology 78: 233.

Harrison, S. 2000. Book review: Savannas, barrens and rock outcrop plant communities of North America, eds. R. C. Anderson, J. S. Fralish and J. M. Baskin. New Phytologist 146:

Amarasekare, P. and S. Harrison 1997. Book review: Population dynamics in ecological space and time, eds. O. R. Rhodes, R. K. Chesser and M. H. Smith; University of Chicago Press. Trends in Ecology and Evolution 12: 78-79.

Harrison, S. 1993. Book review: Risk Assessment in Conservation Biology, by M. Burgman, S. Ferson and H. R. Akcakaya; Chapman and Hall. Trends in Ecology and Evolution 8: 225-6.

Harrison, S. 1990. Book review: Living in a Patchy Environment, B. Shorrocks and I.R. Swingland (eds.), 1989, Cambridge University Press. Nature 348: 123.

Harrison, S. 1988. Book review: Conservation Biology: The Science of Scarcity and Diversity, M.E. Soule (ed.), 1987, Cambridge University Press. Ecology 69: 876-877.

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NSF Research Planning Grant, “Linking Population Processes and Biogeography: A Geographic Study of an Insect Herbivore” June 1993-Dec. 1994 ($18,000).

NSF Conservation Biology, “Plant Communities on Patchy Serpentine Outcrops: Spatial Structure and Ecological Interactions,” June 1995-May 1998 ($150,000).

NSF Ecology, “Spatial Dynamics of an Outbreaking Insect Population” (with A. Hastings), July 1996 - June 1999 ($200,000)

NSF Ecology, "Understanding the regional distribution of native meadow remnants in Northern California," July 1999 - June 2002 ($150,000).

National Center for Ecological Analysis and Synthesis, "Change in California Grasslands: A Workshop Proposal" (with M. Stromberg, C. D'Antonio, C. Malmstrom, and K. Rice), June 2000-June 2002 ($30,000).

NSF Ecology, Small Grants for Exploratory Research (SGER), "Effects of Wildfire on Aquatic and Terrestrial Ecology in an Oak Woodland-Chaparral Ecosystem", January 2000-January 2001 ($50,000).

Andrew W. Mellon Foundation, "Plant Diversity in Spatially Complex Landscapes: Ecological and Evolutionary Responses to Environmental Structure," June 2000-May 2003 (with S. Collinge, R. Grosberg, K. Rice, M. Stanton, D. Thiede), $300,000.

NSF Ecology, "Regional and local diversity in a patchy environment: serpentine plants in California," July 2000-June 2003 ($271,000).

Packard Foundation Interdisciplinary Science Program, “Ecological and Evolutionary Responses to Habitat Mosaics: Integrating Across Spatial and Temporal Hierarchies of Plant Biodiversity.” July 2000- June 2005 (with K. Rice, S. Collinge, P. Gepts, and M. Stanton), $998,200.

NSF Ecology, "LTREB: Adding the temporal component to our understanding of grassland diversity." August 2005-July 2010. $293,000

NSF Ecology, "Climate and Competition as Controls on the Persistence and Richness of an Edaphic Endemic Flora “ (with M. Schwartz, E.Damschen). June 2006 – June 2009. $235,000

National Center for Ecological Analysis and Synthesis, “The Role of Niche Conservatism in Producing Biodiversity Gradients” (with H. Cornell and C. McCain). $56,600. 2008-2010.

NSF Ecology, "Testing a Mechanism for the Productivity-Beta Diversity Relationship in Plants“ (with E.Damschen). July 2010 – June 2013. $339,000

Kearney Foundation of Soil Science, "Climate Change and the Future of California's Serpentine Flora: Using Geologic and Soil Information to Improve Conservation Strategies" (with R. J. Southard and H. W. Day), $120,000. January 1, 2010 – June 30, 2012.

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Panelist, National Science Foundation, Ecology program


Member of Council / Governing Board, Ecological Society of America


Associate Editor, Conservation Biology


Associate Editor, Acta Oecologica


Editorial Board, American Naturalist


Young Investigators Award committee, American Society of Naturalists


Research Advisory Committee, Organization for Tropical Studies


Research Committee, Ecological Society of America


Biological Observatory Network committee, National Science Foundation


Scientific Advisory Board, National Center for Ecological Analysis and Synthesis


Publications committee, Ecological Society of America


Vice-President, American Society of Naturalists


Editorial board, Landscape Ecology

2009- Editorial board, American Naturalist
2010, 2013 National Science Foundation, Ecology Panel
2013 Reviewer, Botany Graduate Program, Oregon State University

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Chair, Conservation Biology Area of Emphasis, Graduate Group in Ecology


Campus Coordinator, UC Natural Reserve System (UCNRS)


Campus Director, UCNRS


 Universitywide Advisory Committee, UCNRS


John Muir Institute of the Environment Steering Committee


 Associate Dean search, College of Agricultural & Environmental Sciences


Committee to review the Scripps Coastal Reserve, UCNRS


Chair, Sedgwick Ranch Reserve planning workshop, UCNRS


 Public Service Research Program Steering Committee


Putah Creek Riparian Reserve Advisory Committee


 Geology Graduate Program review committee


Chair, Biological Invasions IGERT Curriculum Committee


Biological Invasions IGERT Steering Committee


 Chair, Environmental Biology Strategic Planning Committee, College of Agricultural and Environmental Sciences


Committee to review the Fort Ord Reserve, UCNRS


Chair, committee to review the proposed Blue Oak Ranch Reserve, UCNRS


Center for Population Biology Scoping Committee


Chair, Academic and Stretegic Planning Committee, College of Agricultural and Environmental Sciences


Term Appointment Review Committee, CAES


Review Committee, John Muir Institute of the Environment

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Reviewer, “A Multi-Resource Conservation Strategy for the Northern Spotted Owl,” for U.S. Forest Service.


Expert witness, Seattle Audubon Society vs. Moseley, regarding the U.S Forest Service’s management plans for the Northern Spotted Owl.


Reviewer, “Biodiversity and the Great Whale Hydroelectric Project,” for Canada.


Invited speaker, National Stewardship Conference, The Nature Conservancy.


Reviewer, Ecological Society of America policy statements on the Endangered Species Act and the Forest Ecosystem Management Assessment Team (FEMAT) report.


Reviewer, proposed revisions to management plans for the Tongass National Forest, for Association of Forest Service Employees for Environmental Ethics(AFSEEE).


Panelist, Wildlife/Aquatic Community Assessment workshop for California Spotted Owl management plan, for U.S. Forest Service.


Invited speaker, Priority Program on the Environment, Zurich, Switzerland.


Contributing author, Global Biodiversity Assessment, by the Scientific Committee on Problems of the Environment (SCOPE) for United Nations Environmental Program


Reviewer, Draft Environmental Impact Statement by U.S. Forest Service, “Managing California Spotted Owl Habitat in the Sierra Nevada National Forests of California,” for Natural Resources Defense Council


Contributing author, The AFSEEE Alternative for the Tongass National Forest, for Association of Forest Service Employees for Environmental Ethics


Reviewer, Draft translocation guidelines for the Karner Blue butterfly, for the New York State Department of Conservation


Reviewer, Draft Recovery Plan for Mitchell’s Satyr, for US Fish and Wildlife Service


Invited speaker, National Biological Survey, Washington DC


Advisor on the management of the Karner Blue Butterfly, for The Nature Conservancy and the Albany Pine Bush Commission


Member, Board of Advisors, Association of Forest Service Employees for Environmental Ethics


Reviewer, Recovery Plan for Serpentine Soil Species of the SF Bay Area,

for US

Fish and Wildlife Service (USFWS)


Member, Board of Trustees, Land Trust of Napa County


Member, Science Advisory Panel, Land Trust of Napa County


Steering committee, Blue Ridge - Berryessa Natural Area Conservation Partnership


Reviewer, Critical Habitat Designation for the Bay Checkerspot Butterfly, for USFWS


Member, Yreka Phlox Recovery Team, for USFWS


Evaluator, proposed methods for prioritizing species for listing actions under the Endangered Species Act, for US Fish and Wildlife Service.


Reviewer, Recovery Plan for the Karner Blue Butterfly, for USFWS


Contractor, Knoxville Wildlife Area Management Plan, California Department of Fish and Game


Reviewer, Regional management plan, US Bureau of Land Management


Reviewer, Payne Ranch grazing plan, US Bureau of Land Management


Member, Science Advisory Team, Santa Clara County Natural Community Conservation Plan (NCCP)

2008-2010 Robert Louis Stevenson State Park Area Planning Committee, Land Trust, Napa County
2009-2012 Organizing Committee, California Native Plant Society Science Conference
2013 Science Advisory Board, Siskiyou Field Institute

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University of Washington, March 2000

Society for Conservation Biology symposium, "Spatial Issues in Conservation", June 2000

California Botanical Society, UC Berkeley, October 2000

Center for Population Biology, UC Davis, December 2000

Ecological Society of America symposium, August 2001
The Nature Conservancy, Making Advances in Conservation Science Hit the Ground,2001

Archbold Field Station, November 20011

Stanford University, May 2002

Davis Botanical Society, June 2002

Geological Society of America, Coleman Symposium, December 2003

University of Delaware, December 2003

California Academy of Sciences, January 2004

North Carolina State University, Brandt Lectureship, March 2004

Univ. of Utah, Stegner Center for Environmental Law, symposium on wilderness, April 20044

University of California, Los Angeles, May 2004

University of Michigan, First Annual Young Scientists Symposium keynote, October 2004

University of Washington, October 2004

University of Helsinki, symposium on spatial aspects of plant-insect interaction, November 2004

American Society of Naturalists, June 2006

Botanical Society of America, August 2006

University of British Columbia, October 2007

UC Davis, workshop on metabolic theory of ecology, May 2008

UC Irvine, October 2010

Florida State University, October 2010

UC Merced, April 2011

UC Berkeley , April 2011

University of Montana, Missoula, October 2011

University of Colorado, Boulder, November 2011

University of Wisconsin, Madison , December 2011

Potchefstroom University, South Africa, February 2012

Oulo University, Finland, September 2012 (5-day short course)

Georgia Technical University, September 2012

UC Santa Cruz, November 2012

Humboldt State University , December 2012

UC Santa Barbara, April 2013

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